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This creature has flying as long as it's modifiedmodifiedEquipment, Auras you control, and counters are modifications.. (Equipment, Auras you control, and counters are modifications.)
PersistPersistWhen this creature dies, if it had no -1/-1 counters on it, return it to the battlefield under its owner's control with a -1/-1 counter on it. (When this creature dies, if it had no -1/-1 counters on it, return it to the battlefield under its owner's control with a -1/-1 counter on it.)
Hover an underlined rule to see what it means.
Read from the rules text. A card that enables a keyword makes or is the thing; a card that rewards it gets better when you have it. Two cards interact when one enables what the other rewards.